Publications

Evidence of an activity-enhancing conformational shift in Arabidopsis thaliana Plant Cysteine Oxidase 4 induced by binding of substrate or substrate-mimics 

Rebecca Latter, J. C. J. Hintzen, L. M. N. Shah, D. M. Gunawardana, R. M. Sher, M. D. White, J. Mecinovic, J. L. P. Benesch, E. Flashman  

https://doi.org/10.1016/j.jbc.2025.110770

PREPRINT: Engineering reduced activity in the oxygen-sensing Arabidopsis thaliana plant cysteine oxidase 4 enzyme results in improved flood resilience. 

Anna Dirr, Arianna Del Greco, Simon Howe, Leah S. Walker, Monica Perri, Vinay Shukla, Francesco Licausi, Emily Flashman 

https://doi.org/10.1101/2025.07.24.666444

PREPRINT: Manipulating plant oxygen sensing through NCO substitution reveals trade-offs between growth and flooding tolerance

Monica Perri, Yuming He, Ximena Chirinos, Mikel Lavilla-Puetra, Rebecca Latter, Thomas Bancroft, Antonis Papachristodoulou, Emily Flashman, Francesco Licausi 

https://doi.org/10.1101/2025.07.24.665511

Hypoxia-activated fluorescent probes as markers of oxygen levels in plant cells and tissues 

Monica Perri, M Shahneawz Khan, Antoine L D Wallabregue, Viktoriia Voloboeva, Amber M Ridgway, Edward N Smith, Hannah Bolland, Ester M Hammond, Stuart J Conway, Daan A Weits, Emily Flashman 

https://doi.org/10.1111/nph.70202

Oxygen sensing and plant adaptation to flooding in a changing climate 

Tilo Renziehausen, Anna Dirr, Romy Schmidt-Schippers, Emily Flashman, Jos Schippers

https://doi.org/10.1098/rstb.2024.0238

Measuring plant cysteine oxidase interactions with substrates using intrinsic tryptophan fluorescence

Dona M. Gunawardana, Daisy A. Southern, Emily Flashman 

https://www.nature.com/articles/s41598-024-83508-y

N-terminal cysteine acetylation and oxidation patterns may define protein stability 

Karen C. Heathcote, Thomas P. Keeley, Matti Myllykoski, Malin Lundekvam, Nina McTiernan, Salma Akter, Norma Masson, Peter J. Ratcliffe, Thomas Arnesen, Emily Flashman 

https://doi.org/10.1038/s41467-024-49489-2

PREPRINT: H2O2 repurposes the plant oxygen-sensing machinery to control the transcriptional response to oxidative stress

Salma Akter, Monica Perri, Mikel Lvilla-Puerta, Beatrice Ferretti, Laura Dalle Carbonare, Vinay Shukla, Yuri Terlara, Daai Zhang, William K. Myers, Beatrice Giuntoli, Emily Flashman, Francesco Licausi 

https://doi.org/10.1101/2024.10.21.619351

Contrasted NCED gene expression across conifers with rising and peaking abscisic acid responses to drought

Gabriele Rizzuto, Dapeng Wang, Jinhui Chen, Tin Hang Hung, Anna Charlott Fitzky, Emily Flashman, John J. MacKay

https://doi.org/10.1016/j.stress.2024.100574

Destined for destruction: The role of methionine aminopeptidases and plant cysteine oxidases in N-degron formation

Andrea Fuentes-Terron, Rebecca Latter, Samuel Madden, Isabel Manrique-Gil, Jessenia Estrada, Noelia Arteaga, Inmaculada Sanchez-Vicente, Oscar Lorenzo, Emily Flashman 

https://doi.org/10.1093/plphys/kiae667

Kinetic measurements to investigate the oxygen-sensing properties of plant cysteine oxidases 

Anna Dirr, Dona M. Gunawardana, Emily Flashman 

https://doi.org/10.1007/978-1-0716-3080_13

Identification of novel plant cysteine oxidase inhibitors from a yeast chemical genetic screen

Mikel Lavilla-Puerta, Rebecca Latter, Francesca Belle, Tiziana Cervelli, Alvaro Galli, Pierdomenico Perata, Andrea Chini, Emily Flashman, Beatrice Giuntoli 

https://doi.org/10.1016/j.jbc.2023.105366

Targeting plant cysteine oxidase activity for improved submergence tolerance

Leah J. Taylor-Kearney, Emily Flashman 

https://doi.org/10.1111/tpj.15605

Plant Cysteine Oxidase Oxygen-Sensing Function Is Conserved in Early Land Plants and Algae 

Leah J. Taylor-Kearney, Samuel Madden, Jack Wilson, William K. Myers, Dona M. Gunawardana, Elisabete Pires, Philip Holdship, Anthony Tumber, Rosalind E. M. Rickaby, Emily Flashman 

https://pubs.acs.org/doi/full/10.1021/acsbiomedchemau.2c00032

Emerging roles of thiol dioxygenases as oxygen sensors

Dona M. Gunawardana, Karen C. Heathcote, Emily Flashman 

https://doi.org/10.1111/febs.16147

Measuring ROS and redox markers in plant cells 

Salma Akter, Shahneawz M Khan, Edward N Smith, Emily Flashman 

https://doi.org/10.1039/d1c00071c

Structures of Arabidopsis thaliana oxygen-sensing plant cysteine oxidases 4 and 5 enable targeted manipulation of their activity 

Mark D. White, Laura Dalle Carbonare, Mikel Lavilla Puerta, Emily Flashman 

https://doi.org/10.1073/pnas.2000206117

Oxygen-sensing mechanisms across eukaryotic kingdoms and their roles in complex multicellularity

Emma U. Hammarlund, Emily Flashman, Sophie Mohlin, Francesco Licausi

https://doi.org/10.1126/science.aba3512

Hypoxia and hypoxia mimetics differentially modulate histone post-translational modifications 

Kuo-Feng Hsu, Sarah E Wilkins, Richard J Hopkinson, Rok Sekirnik, Emily Flashman, Akane Kawamura, James S O McCullagh, Louise J Walport, Christopher J Schofield

https://doi.org/10.1080/15592294.2020.1786305

Zinc excess induces a hypoxia-like response by inhibiting cysteine oxidases in poplar roots

Laura Dalle Carbonare, Mark D. White, Vinay Shukla, Alessandra Francini, Pierdomenico Perata, Emily Flashman, Luca Sebastiana, Francesco Licausi

https://doi.org/10.1104/pp.18.01458

Conserved N-terminal cysteine dioxygenases transduce responses to hypoxia in animals and plants 

Norma Masson, Thomas P. Keeley, Beatric Giuntoli, Mark D. Whote, Mikel Lavilla Puerta, Pierdomenico Perata, Richard J. Hopkinson, Emily Flashman, Francesco Licausi, Peter J. Ratcliffe

https://doi.org/10.1126/science.aaw0112

The plant cysteine oxidases from Arabidopsis thaliana are kinetically tailored to act as oxygen sensors

Mark D. White, Jos J.A.G. Kamps, Samuel East, Leah J. Taylor-Kearney, Emily Flashman 

https://doi.org/10.1074/jbc.RA118.003496

Oxygen-dependent proteolysis regulates the stability of angiosperm polycomb repressive complex 2 subunit VERNALISATION 2 

Daniel J Gibbs, Hannah M Tedds, Anne-Marie Labandera, Mark Bailey, Mark D White, Sjon Hartman, Colleen Sprigg, Sophie L Mogg, Rory Osborne, Charlene Dambire, Tinne Boeckx, Zachary Paling, Laurentius A C J Voesenek, Emily Flashman, Michael J Holdsworth 

https://www.nature.com/articles/s41467-018-07875-7

Plant Cysteine Oxidases are dioxygenases that directly enable arginyl transferase-catalysed arginylation of N-end rule targets

Mark D. White, Maria Klecker, Richard J. Hopkinson, Daan A. Weits, Carolin Mueller, Christin Naumann, Rebecca O'Neill, James Wickens, Jiayu Yang, Jonathan C. Brooks-Bartlett, Elspeth F. Garman, Tom N. Grossmann, Nico Dissmeyer, Emily Flashman 

https://doi.org/10.1038/ncomms14690

Catalytic strategies of the non-heme iron dependent oxygenases and their roles in plant biology 

Mark D. White, Emily Flashman 

https://doi.org/10.1016/j.cbpa.2016.02.017

How decreasing oxygenase activity helps cells cope with hypoxia 

Emily Flashman 

https://doi.org/10.1042/BIO03805026